Automatic cell-type harmonization and integration across Human Cell Atlas datasets
Cite
Xu, C., Prete, M., Webb, S., Jardine, L., Stewart, B.J., Hoo, R., He, P., Meyer, K.B. and Teichmann, S.A., 2023. Automatic cell-type harmonization and integration across Human Cell Atlas datasets. Cell, 186(26), pp.5876-5891.
https://doi.org/10.1016/j.cell.2023.11.026
Abstract
Harmonizing cell types across the single-cell community and assembling them into a common framework is central to building a standardized Human Cell Atlas. Here, we present CellHint, a predictive clustering tree-based tool to resolve cell-type differences in annotation resolution and technical biases across datasets. CellHint accurately quantifies cell-cell transcriptomic similarities and places cell types into a relationship graph that hierarchically defines shared and unique cell subtypes. Application to multiple immune datasets recapitulates expert-curated annotations. CellHint also reveals underexplored relationships between healthy and diseased lung cell states in eight diseases. Furthermore, we present a workflow for fast cross-dataset integration guided by harmonized cell types and cell hierarchy, which uncovers underappreciated cell types in adult human hippocampus. Finally, we apply CellHint to 12 tissues from 38 datasets, providing a deeply curated cross-tissue database with ∼3.7 million cells and various machine learning models for automatic cell annotation across human tissues.
Datasets
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Sample ID
Age group
Leiden clustering
Litvinukova et al. 2020451,513 cells Tucker et al. 2020287,269 cells Koenig et al. 2022150,582 cells Kuppe et al. 202241,648 cells 01. Fibroblast I39,083 cells 02. Fibroblast II38,555 cells 03. Atrial Cardiomyocyte34,051 cells Cardiomyocytes33,698 cells 04. Ventricular Cardiomyocyte I28,671 cells 05. Cytoplasmic Cardiomyocyte I24,695 cells 06. Ventricular Cardiomyocyte II24,528 cells Cardiomyocyte18,310 cells 08. Macrophage17,468 cells 09. Endothelium I17,142 cells 10. Endothelium II10,781 cells 12. Cytoplasmic Cardiomyocyte II6,070 cells 13. Vascular Smooth Muscle5,740 cells 14. Fibroblast III5,582 cells 15. Ventricular Cardiomyocyte III4,707 cells 17. Lymphocyte1,503 cells Ventricular cardiomyocytes235,692 cells Capillary endothelial cells87,214 cells Ventricle-enriched pericytes83,573 cells Atrial cardiomyocytes57,942 cells Cytoplasmic cardiomyocytes29,409 cells Arterial endothelial cells28,741 cells Atria-enriched endothelial cells21,909 cells Venous endothelial cells16,610 cells Stromal pericytes15,871 cells Atria-enriched pericytes14,592 cells MT-high fibroblasts9,687 cells Neuronal cells7,354 cells CMC-like endothelial cells6,947 cells Immune-related endothelial cells4,914 cells CMC-like pericytes3,381 cells Lymphatic endothelial cells2,766 cells Fibroblast-like endothelial cells2,564 cells Mesothelial cells1,696 cells 10x 3' v2, single-nucleus440,543 cells 10x 3' v3, sinlge-nucleus217,869 cells 10x 5' v1, single-nucleus138,028 cells heart left ventricle369,683 cells left cardiac atrium144,374 cells heart right ventricle128,934 cells right cardiac atrium120,967 cells apex of heart91,748 cells interventricular septum75,306 cells African American40,692 cells HCAHeart784400111,999 cells 1_1_1_1_1_1_1_1_1_1_1_1_1_111,414 cells HCAHeart810286211,091 cells 1_1_1_1_1_1_1_1_110,865 cells 2_1_1_1_1_1_1_1_1_110,679 cells HCAHeart784400410,672 cells HCAHeart81028619,290 cells HCAHeart78439996,172 cells HCAHeart79053286,108 cells HCAHeart78351485,319 cells HCAHeart78299795,241 cells HCAHeart78338555,201 cells HCAHeart78299764,821 cells HCAHeart79053324,808 cells HCAHeart78440034,714 cells HCAHeart79053274,534 cells HCAHeart77028814,339 cells HCAHeart78338544,294 cells HCAHeart78338524,237 cells HCAHeart82871274,183 cells HCAHeart77028824,169 cells HCAHeart77028764,121 cells HCAHeart77028774,056 cells HCAHeart78299784,042 cells HCAHeart76646523,954 cells HCAHeart78889283,884 cells HCAHeart78299773,788 cells HCAHeart82871283,761 cells HCAHeart78808623,704 cells HCAHeart78505513,652 cells HCAHeart78351493,651 cells HCAHeart78440003,632 cells HCAHeart77028733,627 cells HCAHeart78366823,496 cells HCAHeart78366843,424 cells HCAHeart77028803,348 cells HCAHeart76565393,346 cells HCAHeart82871263,341 cells HCAHeart76646543,326 cells HCAHeart76980153,325 cells HCAHeart76646533,323 cells HCAHeart76980163,319 cells HCAHeart77028753,175 cells HCAHeart78889293,173 cells HCAHeart78338533,171 cells HCAHeart77028743,054 cells HCAHeart78889262,952 cells HCAHeart78889232,928 cells HCAHeart78889272,895 cells HCAHeart78366832,877 cells HCAHeart81028682,853 cells HCAHeart79053312,847 cells HCAHeart79645132,718 cells HCAHeart78889222,635 cells HCAHeart78366812,571 cells HCAHeart78505492,478 cells HCAHeart78505462,433 cells HCAHeart81028582,349 cells HCAHeart77459682,263 cells HCAHeart78889252,248 cells HCAHeart78808612,176 cells HCAHeart77028792,168 cells HCAHeart77028782,027 cells HCAHeart78889242,015 cells HCAHeart81028661,987 cells HCAHeart78440021,892 cells HCAHeart78505401,734 cells HCAHeart76565361,730 cells HCAHeart81028651,719 cells HCAHeart77576391,712 cells HCAHeart78505481,564 cells HCAHeart79850861,543 cells HCAHeart81028601,516 cells HCAHeart79053301,452 cells HCAHeart77459691,447 cells HCAHeart79850881,334 cells HCAHeart78808601,317 cells HCAHeart79850871,278 cells HCAHeart81028591,272 cells HCAHeart79850891,169 cells HCAHeart78505471,161 cells HCAHeart81028671,071 cells HCAHeart79053291,037 cells HCAHeart76980171,028 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Age group
Sample ID
Leiden clustering
Siletti et al. 2022345,593 cells Franjic et al. 2022191,529 cells Ayhan et al. 2021129,655 cells Tran et al. 202110,068 cells Oligo OPALIN LAMA232,718 cells Oligo OPALIN SLC5A1129,799 cells DG GC PROX1 SGCZ29,497 cells Astro AQP4 GFAP16,233 cells OPC PDGFRA GRIA415,210 cells Micro C1QB P2RY1212,689 cells Astro AQP4 CHRDL19,687 cells Oligo CPXM2 KANK45,367 cells DG GC PROX1 PDLIM54,470 cells CA3 CFAP299 SYN34,229 cells Oligo OPALIN LINC010983,080 cells Endo CLDN5 VWF3,062 cells CA1 dorsal GRIK1 GRM32,983 cells OPC PDGFRA EGR12,401 cells CA1 ventral ACVR1C SYT132,203 cells Micro C1QB CD832,169 cells InN LAMP5 CHST91,460 cells SUB proximal ROBO1 COL5A21,191 cells vSMC ABCC9 P2RY14674 cells InN NR2F2 SLC17A8649 cells InN SST ADAMTS12639 cells VLMC COL1A1 COL1A2588 cells DG MC ARHGAP24 DLC1444 cells SUB distal FN1 NTNG1432 cells aSMC ACTA2 TAGLN416 cells InN SST EPB41L4A394 cells InN PVALB PLEKHH2381 cells InN NR2F2 MIR4300HG309 cells SUB proximal ROBO1 SEMA3E304 cells COP GPR17 ADAM33294 cells Macro F13A1 COLEC12238 cells InN LHX6 AC008415.1214 cells aEndo DKK2 FBLN5123 cells InN MEIS2 SHISAL2B88 cells Oligodendrocytes196,021 cells SGCZ+ granule cells96,326 cells P2RY12+ microglia35,945 cells GFAP high astrocytes35,574 cells CA ACVR1C ExN31,529 cells GFAP low astrocytes30,138 cells CA GLYATL1 ExN15,241 cells PDLIM5+ granule cells13,669 cells CA/Sub COBLL1 ExN12,763 cells CA/Sub GFRA1 ExN9,579 cells CA/Sub CUX2 ExN9,240 cells LAMP5 CHST9 InN8,837 cells CD83+ microglia8,436 cells CA/Sub TRPC3 ExN5,118 cells CA/Sub PXDN ExN4,160 cells Endothelial cells4,064 cells CA/Sub KCNK13 ExN3,170 cells LAMP5 NMBR InN2,935 cells Ependymal cells1,913 cells MEIS2 SHISAL2B InN652 cells 10x 3' v3, sinlge-nucleus634,831 cells 10x 3' v2, sinlge-nucleus42,014 cells hippocampal formation676,845 cells African American11,289 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Age group
Sample ID
Leiden clustering
Ren et al. 2021164,945 cells Stephenson et al. 202197,008 cells Yoshida et al. 202146,964 cells Dominguez Conde et al. 202226,999 cells T_CD4_c01-LEF120,518 cells T_CD8_c01-LEF113,237 cells Mono_c3-CD14-VCAN11,495 cells T_CD8_c05-ZNF68310,620 cells T_CD8_c07-TYROBP10,575 cells CD83_CD14_mono6,772 cells Mono_c2-CD14-HLA-DPB16,768 cells T_CD8_c03-GZMK6,759 cells T_CD8_c08-IL2RB6,344 cells Classical monocytes5,850 cells B_c02-MS4A1-CD274,742 cells T_CD4_c04-ANXA24,429 cells T_CD8_c02-GPR1834,370 cells Teffector/EM_CD44,331 cells T_gdT_c14-TRDV24,323 cells T_CD8_c09-SLC4A103,773 cells B_c03-CD27-AIM23,733 cells T_CD4_c02-AQP33,311 cells T_CD8_c04-COTL13,139 cells T_CD4_c11-GNLY2,475 cells Mono_c1-CD14-CCL32,473 cells T_CD4_c03-ITGA42,328 cells T_CD4_c12-FOXP32,250 cells T_CD4_c06-NR4A22,233 cells T_CD4_c09-GZMK-FOS_l1,908 cells T_CD4_c07-AHNAK1,841 cells B_switched_memory1,049 cells B_c04-SOX5-TNFRSF1B838 cells T_CD4_c08-GZMK-FOS_h815 cells Mono_c4-CD14-CD16662 cells T_CD8_c10-MKI67-GZMK649 cells Monocyte CD14 IFN stim551 cells B_non-switched_memory499 cells NK_CD56bright_CD16-445 cells Nonclassical monocytes236 cells T_CD8_c13-HAVCR2171 cells T_CD4_c13-MKI67-CCL5_l126 cells T CD4 naive IFN stim117 cells T_CD8_c12-MKI67-TYROBP94 cells Monocyte CD16 IFN stim90 cells Monocyte CD14 IL616 cells Tcm/Naive helper T cells73,640 cells CD16+ NK cells47,171 cells Classical monocytes44,292 cells Tem/Temra cytotoxic T cells34,734 cells Tem/Effector helper T cells28,005 cells Tem/Trm cytotoxic T cells17,351 cells Tcm/Naive cytotoxic T cells16,755 cells Naive B cells16,695 cells Non-classical monocytes8,642 cells Switched memory B cells5,558 cells Non-switched memory B cells5,285 cells Regulatory T cells3,291 cells Tem/Temra helper T cells2,758 cells Megakaryocytes/platelets2,710 cells Memory B cells2,216 cells Age-associated B cells1,163 cells CD16- NK cells1,016 cells Cycling immune mix863 cells C1 non-classical monocytes238 cells late adult stage8,130 cells CV001_KM102023958,439 cells CV001_KM102023988,176 cells CV001_KM102023997,277 cells CV001_KM102024006,620 cells CV001_KM102024016,455 cells CV001_KM102023944,977 cells CV001_KM102023972,121 cells CV001_KM102023961,984 cells CV001_KM10202418639 cells CV001_KM10202419276 cells CZI-IA10466284+CZI-IA10466380204 cells CZI-IA10466281+CZI-IA10466377191 cells CZI-IA10466283+CZI-IA10466379189 cells CZI-IA10466282+CZI-IA10466378183 cells CZI-IA10244331+CZI-IA10244433103 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Age group
Sample ID
Leiden clustering
Madissoon et al. 2022105,135 cells Adams et al. 202095,301 cells Travaglini et al. 202060,971 cells Madissoon et al. 202057,019 cells Macrophage_Alveolar23,905 cells Endothelia_vascular_Cap_g7,788 cells Alveolar_Type24,676 cells Macrophage_MARCOpos4,512 cells Capillary Aerocyte4,352 cells Natural Killer4,250 cells Alveolar Epithelial Type 23,814 cells CD4_EM/Effector3,606 cells Fibro_adventitial3,589 cells Fibro_alveolar2,790 cells Endothelia_Lymphatic2,662 cells Endothelia_vascular_Cap_a2,458 cells CD4+ Memory/Effector T2,419 cells Macrophage_intermediate2,309 cells Endothelia_vascular_venous_pulmonary1,762 cells Endothelia_vascular_arterial_pulmonary1,692 cells Endothelia_vascular_venous_systemic1,529 cells Basophil/Mast 11,349 cells Alveolar Fibroblast1,296 cells Muscle_smooth_pulmonary1,159 cells Muscle_pericyte_parenchyma1,087 cells Classical Monocyte1,013 cells Alveolar Epithelial Type 1972 cells Macrophage_MARCOneg937 cells CD8+ Memory/Effector T771 cells Airway Smooth Muscle693 cells Signaling Alveolar Epithelial Type 2669 cells Capillary Intermediate 1628 cells Nonclassical Monocyte604 cells Adventitial Fibroblast557 cells Fibro_myofibroblast538 cells Vascular Smooth Muscle469 cells Capillary Intermediate 2464 cells Secretory_Goblet461 cells Bronchial Vessel 1455 cells Natural Killer T335 cells Macro_alveolar_metallothioneins332 cells IGSF21+ Dendritic278 cells Muscle_pericyte_airway265 cells Fibro_peribronchial258 cells Macro_interstitial251 cells Differentiating Basal249 cells DC_Monocyte_Dividing242 cells Myeloid Dendritic Type 2236 cells Bronchial Vessel 2235 cells Proliferating Macrophage226 cells OLR1+ Classical Monocyte207 cells Intermediate Monocyte188 cells TREM2+ Dendritic159 cells Myeloid Dendritic Type 1130 cells Plasmacytoid Dendritic116 cells Proliferating NK/T101 cells Macrophage_Dividing94 cells Muscle_smooth_syst_arterial92 cells Proximal Ciliated88 cells Endothelia_vascular_arterial_systemic83 cells Fibro_immune_recruiting51 cells Proliferating Basal45 cells Muscle_perivascular_immune_recruiting25 cells Alveolar macrophages58,974 cells CD16+ NK cells34,063 cells Classical monocytes18,806 cells Capillary endothelial cells17,696 cells Tem/Effector helper T cells12,037 cells Intermediate macrophages9,158 cells Adventitial fibroblasts8,374 cells Capillary aerocytes7,492 cells Tem/Trm cytotoxic T cells6,854 cells Tem/Temra cytotoxic T cells6,595 cells Alveolar fibroblasts5,314 cells Ciliated cells4,869 cells Non-classical monocytes4,699 cells Lymphatic endothelial cells4,433 cells Pulmonary venous endothelial cells4,257 cells Arterial endothelial cells3,838 cells Memory B cells3,384 cells Tcm/Naive helper T cells3,174 cells Trm cytotoxic T cells2,742 cells Regulatory T cells2,389 cells Systemic venous endothelial cells2,082 cells CD16- NK cells2,020 cells Myofibroblasts1,312 cells Follicular helper T cells698 cells Suprabasal cells515 cells Cycling macrophages341 cells Mesothelial cells163 cells 10x 5' transcription profiling87,445 cells lung parenchyma57,019 cells lower lobe of left lung44,212 cells upper lobe of left lung25,478 cells Hispanic or Latin American4,315 cells WSSS8062679-010,939 cells HCATisStab77471985,110 cells HCATisStab76460334,361 cells HCATisStab76460324,252 cells HCATisStab75872023,772 cells HCATisStab75097363,718 cells HCATisStab75872083,689 cells HCATisStab77471973,580 cells HCATisStab75872053,471 cells HCATisStab77472003,445 cells WTDAtest7732269-03,424 cells HCATisStab7732262-03,322 cells HCATisStab76599693,299 cells WTDAtest7732270-03,123 cells HCATisStab76460353,037 cells WSSS_A_LNG8620063-02,791 cells HCATisStab77471992,703 cells HCATisStab75872112,631 cells 5841STDY7991475-02,578 cells HCATisStab75097342,347 cells WSSS_A_LNG8620059-02,322 cells HCATisStab7732264-02,303 cells WTDAtest7732268-02,237 cells HCATisStab76599702,050 cells HCATisStab7732263-01,979 cells 5841STDY7991482-01,973 cells HCATisStab75097351,918 cells HCATisStab76460341,713 cells 5841STDY7991479-01,528 cells 5841STDY7991487-01,454 cells HCATisStab76599711,418 cells 5841STDY7991486-01,410 cells 5841STDY7991478-01,294 cells WSSS_A_LNG8620060-01,081 cells WSSS_A_LNG8620068-01,021 cells 5841STDY7991484-0982 cells HCATisStab7732261-0826 cells WSSS_A_LNG8620064-0792 cells 5841STDY7991477-0777 cells 5841STDY7991485-0717 cells WSSS_A_LNG8620065-0588 cells WSSS_A_LNG8620066-0556 cells HCATisStab7659968505 cells 5841STDY7991476-0445 cells WSSS_A_LNG8620067-0304 cells WTDAtest7888000-0243 cells WTDAtest7887999-0233 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Age group
Leiden clustering
Sample ID
Elmentaite et al. 2021113,121 cells Smillie et al. 2019109,884 cells James et al. 202027,680 cells Burclaff et al. 202212,590 cells IgA plasma cell14,494 cells B cell IgA Plasma12,472 cells Enterocyte Progenitors5,319 cells Immature Enterocytes 24,447 cells Immature Enterocytes 14,401 cells BEST4+ epithelial3,822 cells Transitional Stromal 3 (C3+)3,638 cells CD4+ Activated Fos-lo3,559 cells CD4+ Activated Fos-hi3,524 cells BEST2+ Goblet cell3,007 cells Immature Goblet2,886 cells Activated CD8 T2,256 cells WNT2B+ Fos-lo 11,871 cells Activated CD4 T1,704 cells Mature venous EC1,673 cells Best4+ Enterocytes1,292 cells Mature arterial EC1,224 cells Contractile pericyte (PLN+)1,028 cells Follicular B cell754 cells Stromal 1 (CCL11+)573 cells arterial capillary439 cells LYVE1+ Macrophage420 cells Post-capillary Venules345 cells B cell IgG Plasma328 cells Stromal 2 (NPY+)226 cells Cycling Monocytes191 cells Stromal 1 (ADAMDEC1+)165 cells mLN Stroma (FMO2+)129 cells Inflammatory Monocytes123 cells myofibroblast (RSPO2+)114 cells Mesothelium (PRG4+)25 cells Inflammatory Fibroblasts20 cells Trm cytotoxic T cells12,488 cells Cycling/Secretory TA7,345 cells Type 17 helper T cells5,708 cells BEST4+ enterocytes4,903 cells gamma-delta T cells4,006 cells Follicular B cells3,790 cells Type 1 helper T cells3,115 cells Tcm/Naive helper T cells3,093 cells Immature goblet cells2,708 cells Regulatory T cells2,619 cells Memory B cells2,584 cells Tem/Trm cytotoxic T cells2,173 cells Venous endothelial cells2,039 cells Tem/Effector helper T cells1,584 cells Arterial endothelial cells1,279 cells Capillary endothelial cells1,114 cells Lymphatic endothelial cells927 cells Intestinal macrophages807 cells Follicular helper T cells527 cells Germinal center B cells227 cells Enteroendocrine cells213 cells colonic epithelium109,884 cells transverse colon24,585 cells large intestine22,538 cells sigmoid colon18,523 cells ascending colon8,748 cells vermiform appendix6,375 cells descending colon5,142 cells African American3,795 cells WTDAtest78440189,485 cells WTDAtest78440168,596 cells Human_colon_16S80019036,744 cells Human_colon_16S80005136,342 cells WTDAtest78440155,785 cells Human_colon_16S80018794,754 cells WTDAtest78440174,387 cells WTDAtest78440134,147 cells Human_colon_16S80018733,512 cells Human_colon_16S80018653,233 cells 4861STDY71359122,982 cells Human_colon_16S81178312,602 cells 4861STDY71359162,510 cells Human_colon_16S81239112,437 cells 4861STDY72084172,380 cells Human_colon_16S81178302,375 cells 4861STDY71359182,374 cells Human_colon_16S81239172,273 cells 4861STDY71359112,233 cells Human_colon_16S80018692,232 cells Human_colon_16S81178292,164 cells WTDAtest78440242,112 cells 4861STDY72084111,998 cells Human_colon_16S80018851,997 cells Human_colon_16S72556821,942 cells Human_colon_16S81591911,806 cells Human_colon_16S80018671,662 cells WTDAtest78440291,590 cells WTDAtest77707191,569 cells 4861STDY72084101,530 cells Human_colon_16S80018631,477 cells 4861STDY72084161,386 cells Human_colon_16S80026291,353 cells Human_colon_16S81239161,322 cells Human_colon_16S72556761,268 cells 4861STDY72084151,216 cells Human_colon_16S81591931,211 cells 4861STDY71359171,179 cells Human_colon_16S80026261,169 cells WTDAtest77707161,165 cells WTDAtest77707181,162 cells Human_colon_16S80019071,157 cells Human_colon_16S80018711,151 cells Human_colon_16S80026271,024 cells Human_colon_16S81239081,009 cells Human_colon_16S8001883992 cells Human_colon_16S8002630985 cells Human_colon_16S7255681915 cells Human_colon_16S8002628888 cells Human_colon_16S8002624882 cells Human_colon_16S7255680807 cells Human_colon_16S8002623766 cells Human_colon_16S8001881698 cells Human_colon_16S8001905671 cells Human_colon_16S8123920620 cells Human_colon_16S8000511618 cells Human_colon_16S8001878612 cells Human_colon_16S8123913605 cells Human_colon_16S8000471572 cells Human_colon_16S8000515564 cells Human_colon_16S8123919540 cells Human_colon_16S8159192537 cells Human_colon_16S8000481533 cells Human_colon_16S8001871-1533 cells Human_colon_16S8002582510 cells Human_colon_16S8000475451 cells Human_colon_16S8123912445 cells Human_colon_16S8000477394 cells Human_colon_16S8123910381 cells Human_colon_16S8000493377 cells Human_colon_16S8002625363 cells Human_colon_16S8000491346 cells Human_colon_16S8123909321 cells Human_colon_16S8000487319 cells Human_colon_16S8123918291 cells Human_colon_16S8002581264 cells Human_colon_16S8001875259 cells Human_colon_16S8000473254 cells Human_colon_16S8000479252 cells Human_colon_16S8123915249 cells Human_colon_16S8000483208 cells Human_colon_16S8000489194 cells Human_colon_16S7255675127 cells Human_colon_16S8001867-1111 cells Human_colon_16S7255679110 cells Human_colon_16S8001863-175 cells Human_colon_16S800256616 cells Human_colon_16S80025733 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Age
Leiden clustering
Age group
Sample ID
Guilliams et al. 2022167,587 cells Andrews et al. 202273,295 cells Aizarani et al. 201910,352 cells MacParland et al. 20188,444 cells Mono+mono derived cells29,639 cells Circulating NK/NKT12,567 cells Endothelial cells2,834 cells alpha-beta_T_Cells961 cells Inflammatory_Macrophage813 cells gamma-delta_T_Cells_1464 cells Non-inflammatory_Macrophage379 cells Periportal_LSECs327 cells Central_venous_LSECs306 cells Portal_endothelial_Cells211 cells gamma-delta_T_Cells_2105 cells Hepatic_Stellate_Cells37 cells Tem/Trm cytotoxic T cells39,056 cells Periportal hepatocytes34,031 cells Pericentral hepatocytes27,012 cells Classical monocytes18,034 cells Tem/Temra cytotoxic T cells12,662 cells CD16+ NK cells11,831 cells CD16- NK cells11,492 cells Non-classical monocytes11,186 cells Central venous LSECs10,747 cells Interzonal hepatocytes10,200 cells Inflammatory macrophages8,426 cells Non-inflammatory macrophages3,976 cells Cholangiocytes3,631 cells Stellate cells1,997 cells Periportal LSECs1,153 cells Portal endothelial cells747 cells 10x 3' v3, single-nucleus58,915 cells caudate lobe of liver73,295 cells prime adult stage53,022 cells young adult stage20,273 cells prime adult stage53,022 cells young adult stage20,273 cells CD45negASGR1negBio66 cells CD45negASGR1neg31566 cells ASGR1_CD45_UNB_P452 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Leiden clustering
Age group
Sample ID
Perez et al. 2022143,051 cells Tabula Sapiens 202230,734 cells Micheli et al. 202022,058 cells He et al. 20205,732 cells Fast skeletal muscle65,519 cells Slow skeletal muscle47,892 cells mesenchymal stem cell14,148 cells skeletal muscle satellite stem cell4,831 cells T/NK/Macrophages4,567 cells endothelial cell of vascular tree3,602 cells DCN+ GSN+ MYOC+ Fibroblasts3,489 cells COL1A1+ Fibroblasts2,744 cells FBN1+ MFAP5+ CD55+ Fibroblasts2,626 cells capillary endothelial cell2,015 cells CD36+ VWF+ Platelets1,904 cells PAX7+ DLK1+ MuSCs and progenitors1,724 cells ACTA2+ MYH11+ MYL9+ Smooth muscle cells1,662 cells PAX7low MYF5+ MuSCs and progenitors1,468 cells IL7R+ PTPRC+ NKG7+ B/T/NK cells1,319 cells APOD+ CFD+ PLAC9+ Adipocytes1,182 cells Satellite Cell NPPC_high Muscle1,139 cells Satellite Cell APOC1_high Muscle946 cells Endothelial Cell IFI27_high Muscle928 cells ICAM1+ SELE+ VCAM1+ Endothelial875 cells Fibroblast Muscle867 cells CLDN5+ PECAM1+ Endothelial831 cells RGS5+ MYL9+ Pericytes655 cells S100A9+ LYZ+ Inflammatory macrophages647 cells C1QA+ CD74+ Macrophages614 cells cd8-positive, alpha-beta t cell536 cells NK/T Cell Muscle510 cells Smooth Muscle Cell RGS5_high Muscle463 cells fast muscle cell266 cells ACTA1+ Mature skeletal muscle256 cells Endothelial Cell ACKR1_high Muscle228 cells Satellite Cell DLK1_high Muscle222 cells cd4-positive, alpha-beta t cell221 cells Macrophage/Monocyte Muscle175 cells slow muscle cell163 cells Smooth Muscle Cell TAGLN_high Muscle149 cells mature nk t cell137 cells endothelial cell of lymphatic vessel134 cells endothelial cell of artery133 cells HBA1+ Erythroblasts62 cells smooth muscle cell58 cells Fast skeletal muscle61,815 cells Slow skeletal muscle44,450 cells Mesenchymal stem cells28,275 cells Capillary endothelial cells11,215 cells PAX7 low MuSCs7,250 cells PAX7 high MuSCs6,281 cells Ribosome high skeletal muscle4,732 cells Venous endothelial cells4,727 cells Lymphoid/Macrophage4,633 cells LRRK2+ fast skeletal muscle2,613 cells C1QA+ CD74+ macrophages2,132 cells S100A9+ LYZ+ macrophages1,232 cells Arterial endothelial cells130 cells Lymphatic endothelial cells128 cells 10x 5' v1, sinlge-nucleus143,051 cells 10x 5' transcription profiling5,732 cells skeletal muscle organ, vertebrate170,841 cells rectus abdominis muscle11,897 cells muscle of pelvic diaphragm7,814 cells muscle of abdomen6,142 cells African American or Afro-Caribbean18,591 cells TSP14_Muscle_Abdomen_10X_1_16,142 cells TSP2_Muscle_rectusabdominus_10X_1_15,561 cells TSP2_Muscle_rectusabdominus_10X_1_25,117 cells TSP2_Muscle_diaphragm_10X_1_22,181 cells TSP2_Muscle_diaphragm_10X_1_11,938 cells TSP14_Muscle_Diaphragm_10X_1_21,168 cells TSP14_Muscle_Diaphragm_10X_1_11,073 cells TSP4_Muscle_diaphragm_SS2_B112866_B134045301 cells TSP2_Muscle_rectusabdominus_SS2_B113467_B111654298 cells TSP4_Muscle_rectusabdominus_SS2_B114867_B134050291 cells TSP4_Muscle_diaphragm_SS2_B114833_B134044284 cells TSP2_Muscle_diaphragm_SS2_B114616_B134456262 cells TSP4_Muscle_diaphragm_SS2_B112813_B134043261 cells TSP4_Muscle_diaphragm_SS2_B112816_B134051251 cells TSP2_Muscle_diaphragm_SS2_B114580_B104855249 cells TSP4_Muscle_rectusabdominus_SS2_B114953_B134040248 cells TSP4_Muscle_rectusabdominus_SS2_B112802_B134053240 cells TSP2_Muscle_diaphragm_SS2_B114671_B133329207 cells TSP2_Muscle_rectusabdominus_SS2_B113693_B111573185 cells TSP2_Muscle_rectusabdominus_SS2_B113450_B104857182 cells TSP4_Muscle_rectusabdominus_SS2_B114996_B134042172 cells TSP4_Muscle_rectusabdominus_SS2_B114997_B134049162 cells TSP2_Muscle_rectusabdominus_SS2_B114661_B133087106 cells TSP2_Muscle_diaphragm_SS2_B114670_B13308861 cells TSP4_Muscle_diaphragm_SS2_B114835_B13405234 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Leiden clustering
Age group
Sample ID
Madissoon et al. 202092,049 cells Dominguez Conde et al. 202270,099 cells Tabula Sapiens 202234,004 cells He et al. 20204,512 cells Memory B cells13,962 cells Classical monocytes5,714 cells T_CD8_activated5,009 cells classical monocyte4,443 cells cd8-positive, alpha-beta memory t cell4,301 cells NK_CD56bright_CD16-3,891 cells Teffector/EM_CD43,837 cells cd4-positive, alpha-beta memory t cell1,771 cells naive thymus-derived cd4-positive, alpha-beta t cell1,261 cells Erythrophagocytic macrophages1,071 cells CD4 T Cell Spleen953 cells B Cell MS4A1_high Spleen670 cells naive thymus-derived cd8-positive, alpha-beta t cell610 cells endothelial cell596 cells CD8 T Cell GZMK_high Spleen578 cells B Cell TCL1A_high Spleen525 cells mature nk t cell426 cells NK/T Cell Spleen333 cells CD8 T Cell RP11-291B21.2_high Spleen325 cells Macrophage/Monocyte Spleen319 cells Tnaive/CM_CD4_activated306 cells Plasma Cell IGHG1_high Spleen304 cells regulatory t cell288 cells Nonclassical monocytes254 cells CD8 T Cell TRBV4-2_high Spleen158 cells intermediate monocyte131 cells T GD Cell Spleen122 cells innate lymphoid cell117 cells Hematopoietic stem cell73 cells cd141-positive myeloid dendritic cell51 cells cd1c-positive myeloid dendritic cell51 cells plasmacytoid dendritic cell43 cells cd8-positive, alpha-beta t cell43 cells Alveolar macrophages38 cells Plasma Cell IGKV2-30_high Spleen29 cells Intestinal macrophages21 cells Memory B cells35,708 cells Naive B cells26,154 cells Tcm/Naive helper T cells18,607 cells Tem/Trm cytotoxic T cells15,078 cells Classical monocytes12,237 cells CD16+ NK cells11,991 cells CD16- NK cells11,831 cells Follicular helper T cells7,224 cells Tem/Effector helper T cells6,531 cells CRTAM+ gamma-delta T cells4,634 cells Tem/Temra cytotoxic T cells4,212 cells Regulatory T cells4,010 cells Low-quality B cells3,402 cells Erythrophagocytic macrophages3,024 cells Tcm/Naive cytotoxic T cells2,472 cells Cycling immune mix1,973 cells Non-classical monocytes1,557 cells Trm cytotoxic T cells1,322 cells Endothelial cells591 cells Megakaryocytes/platelets82 cells 10x 5' transcription profiling4,512 cells African American or Afro-Caribbean7,679 cells HCATisStab758720321,701 cells HCATisStabAug17727639416,004 cells HCATisStab75872097,234 cells HCATisStab75872006,349 cells CZI-IA9924320+CZI-IA99243686,269 cells TSP14_Spleen_NA_10X_1_15,697 cells TSP14_Spleen_NA_10X_2_15,238 cells HCATisStab74638494,710 cells TSP7_Spleen_NA_10X_1_14,692 cells TSP14_Spleen_NA_10X_1_1_5Prime4,495 cells CZI-IA10244332+CZI-IA102444344,451 cells TSP2_Spleen_NA_10X_1_14,396 cells HCATisStabAug1772763913,993 cells HCATisStab74638463,578 cells TSP14_Spleen_NA_10X_2_1_5Prime3,534 cells HCATisStab74638473,480 cells HCATisStab74638483,456 cells HCATisStabAug1772763923,438 cells HCATisStab75872063,375 cells HCATisStabAug1773765632,562 cells TSP2_Spleen_NA_10X_2_12,302 cells CZI-IA10471909+CZI-IA104720052,295 cells CZI-IA10471910+CZI-IA104720062,282 cells HCATisStabAug1770780162,058 cells HCATisStabAug1770780182,049 cells HCATisStabAug1773765611,900 cells HCATisStabAug1770780191,806 cells HCATisStabAug1773765651,769 cells TSP7_Spleen_NA_10X_2_11,502 cells CZI-IA10466284+CZI-IA104663801,407 cells HCATisStabAug1770780171,406 cells CZI-IA10466281+CZI-IA104663771,294 cells CZI-IA10466283+CZI-IA104663791,266 cells CZI-IA10466282+CZI-IA104663781,243 cells HCATisStabAug1773765671,181 cells TSP7_Spleen_NA_SS2_B134152__ImmuneLinPos356 cells TSP2_Spleen_NA_SS2_B113701_B104856339 cells TSP7_Spleen_NA_SS2_B134167_B114748317 cells TSP7_Spleen_NA_SS2_B134166_B114749283 cells TSP2_Spleen_NA_SS2_B114615_B133086273 cells TSP2_Spleen_NA_SS2_B113451_B133325251 cells TSP7_Spleen_NA_SS2_B134153_B114729211 cells TSP2_Spleen_NA_SS2_B114583_B104854118 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Leiden clustering
Age group
Sample ID
Lake et al. 2021105,428 cells HCA Kidney 202248,772 cells Stewart et al. 201920,319 cells Muto et al. 202119,985 cells PRECISE-9437765,861 cells PRECISE-9454202,437 cells PRECISE165536-000-003N1,611 cells Sample1153-EO31,334 cells Sample1162-EO21,329 cells Sample1157-EO21,166 cells Sample1157-EO11,042 cells Proximal tubule10,202 cells Peritubular capillary endothelium 1653 cells Glomerular endothelium596 cells MNP-a/classical monocyte derived340 cells Thick ascending limb of Loop of Henle302 cells MNP-b/non-classical monocyte derived297 cells Ascending vasa recta endothelium241 cells Type A intercalated cell193 cells Epithelial progenitor cell188 cells MNP-c/dendritic cell172 cells Peritubular capillary endothelium 2163 cells Descending vasa recta endothelium163 cells Type B intercalated cell103 cells MNP-d/Tissue macrophage73 cells Connecting tubule63 cells Distinct proximal tubule 162 cells Proliferating Proximal Tubule57 cells Distinct proximal tubule 257 cells Indistinct intercalated cell24 cells Plasmacytoid dendritic cell18 cells Non-classical monocytes1,093 cells Classical monocytes534 cells 10x 3' v3, sinlge-nucleus85,194 cells 10x 5' v1, single-nucleus19,985 cells cortex of kidney127,417 cells renal papilla20,336 cells renal medulla13,649 cells African American4,234 cells Hispanic or Latin American2,920 cells PreciseHCA54ref4,715 cells CZIKidney76328033,016 cells PreciseHCA51ref2,901 cells CZIKidney76328042,744 cells PreciseHCA55ref1,786 cells 4602STDY69308531,689 cells PRECISE165536000003N1,611 cells PreciseHCA29ref1,603 cells 4602STDY69308521,601 cells LDSample1153EO31,334 cells LDSample1162EO21,329 cells LDSample1157EO21,166 cells 4602STDY69764281,114 cells 4602STDY69764271,079 cells 4602STDY69764261,076 cells LDSample1157EO11,042 cells CZIKidney7587406491 cells CZIKidney7587408430 cells CZIKidney7587407422 cells CZIKidney7587405319 cells CZIKidney7587422101 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Leiden clustering
Age group
Sample ID
Dominguez Conde et al. 202282,974 cells Szabo et al. 201916,527 cells James et al. 202013,872 cells Elmentaite et al. 202111,067 cells Tnaive/CM_CD417,529 cells Memory B cells13,459 cells Teffector/EM_CD47,608 cells Tnaive/CM_CD4_activated3,237 cells Activated CD4 T2,590 cells Follicular B cell1,828 cells NK_CD56bright_CD16-1,367 cells Erythrophagocytic macrophages288 cells mLN Stroma (FMO2+)226 cells Mature venous EC182 cells Classical monocytes143 cells Mature arterial EC118 cells Alveolar macrophages114 cells arterial capillary89 cells B cell IgA Plasma50 cells LYVE1+ Macrophage24 cells Intestinal macrophages21 cells Nonclassical monocytes20 cells Transitional Stromal 3 (C3+)19 cells Tcm/Naive helper T cells37,025 cells Memory B cells17,290 cells Regulatory T cells13,596 cells Follicular helper T cells11,986 cells Activated CD4 T cells9,383 cells Tem/Trm cytotoxic T cells6,996 cells Tcm/Naive cytotoxic T cells4,795 cells Tem/Effector helper T cells3,368 cells CD16- NK cells1,647 cells BTAF1 high T cells1,545 cells BTAF1 high B cells1,070 cells Trm cytotoxic T cells556 cells Erythrophagocytic macrophages413 cells Lymphatic endothelial cells400 cells Endothelial cells389 cells Germinal center B cells348 cells mLN Stroma (FMO2+)225 cells Age-associated B cells164 cells Classical monocytes119 cells Non-classical monocytes76 cells thoracic lymph node60,530 cells mesenteric lymph node46,600 cells bronchopulmonary lymph node16,527 cells Hispanic or Latin American8,345 cells Human_colon_16S81239215,613 cells Donor2_lymph_node_act5,190 cells CZI-IA10471910+CZI-IA104720065,169 cells CZI-IA10244332+CZI-IA102444345,123 cells CZI-IA10471909+CZI-IA104720054,486 cells Donor1_lymph_node_rest4,191 cells Donor1_lymph_node_act4,154 cells CZI-IA9924321+CZI-IA99243693,637 cells CZI-IA10466284+CZI-IA104663803,484 cells CZI-IA10466282+CZI-IA104663783,224 cells CZI-IA10466281+CZI-IA104663773,100 cells 4861STDY71359143,086 cells CZI-IA10466283+CZI-IA104663793,031 cells Donor2_lymph_node_rest2,992 cells 4861STDY71359132,975 cells Human_colon_16S81178282,949 cells Human_colon_16S80004841,276 cells 4861STDY72084131,194 cells 4861STDY72084121,117 cells Human_colon_16S72556781,088 cells Human_colon_16S8000484-1966 cells Human_colon_16S7255677646 cells Human_colon_16S8159194537 cells CZI-IA9924320+CZI-IA9924368416 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Leiden clustering
Age group
Sample ID
Tosti et al. 202179,367 cells Tritschler et al. 202226,474 cells Fasolino et al. 202213,960 cells Muraro et al. 20162,115 cells alpha_stress II3,715 cells alpha_immature1,961 cells Activated Stellate1,609 cells beta_stress II1,259 cells duct_acinar_related762 cells beta_MHC/autoantigen349 cells beta_mtDNA deficient274 cells Quiescent Stellate143 cells acinar_minor_mhcclassII125 cells immune_stellates115 cells Idling acinar cells35,308 cells Secretory acinar cells21,444 cells Stressed beta cells2,565 cells Endothelial cells2,106 cells Activated stellate cells2,066 cells Quiescent stellate cells142 cells MUC5B+ ductal cells62 cells 10x 3' v2, single-nucleus79,367 cells islet of Langerhans40,434 cells 
Data source
Subject ID
Cell subtype
CellHint harmonised group
Cell type
Assay
Disease
Sex
Tissue
Ethnicity
Age
Leiden clustering
Age group
Sample ID
Dominguez Conde et al. 202239,889 cells Tabula Sapiens 202212,293 cells Roy et al. 202111,201 cells He et al. 20203,230 cells Classical monocytes5,032 cells Teffector/EM_CD42,637 cells CD24 Neutrophil2,337 cells Myeloid 1 (neutrophil)1,387 cells Early lymphoid1,016 cells NK_CD56bright_CD16-832 cells CD4 T Cell IL7R_high Marrow792 cells erythroid progenitor757 cells Megakaryocyte/Erythroid Progenitor647 cells hematopoietic stem cell617 cells CD8 T Cell RGS1_high Marrow612 cells Nonclassical monocytes586 cells Erythroid (cycling)463 cells NAMPT neutrophil443 cells Plasmacytoid dendritic precursor419 cells Myeloid 2 (cycling)403 cells Myeloid 2 (monocytic)402 cells myeloid progenitor287 cells B lymphoid (cycling)286 cells NK/T Cell GNLY_high Marrow242 cells Monocyte/Dendritic precursor (cycling)238 cells NK/T Cell PTGDS_high Marrow201 cells Megakaryocyte/Erythroid (cycling)201 cells CD8 T Cell TRBV4-2_high Marrow197 cells Tnaive/CM_CD4_activated180 cells Eosinophil/Basophil/Mast171 cells Macrophage/Monocyte Marrow145 cells T GD Cell TRDV1_high Marrow124 cells Plasmacytoid dendritic precursor (cycling)114 cells CD8 T Cell TRBV4-1_high Marrow89 cells Multiple Progenitor Marrow54 cells T GD Cell TRDV2_high Marrow42 cells Plasma Cell JCHAIN_high Marrow42 cells High Proliferation Erythrocyte Marrow42 cells Erythrophagocytic macrophages36 cells CD8 T Cell TRAV14DV4_high Marrow26 cells Intermediate macrophages25 cells Alveolar macrophages19 cells CD16+ NK cells8,779 cells Tcm/Naive helper T cells7,982 cells Classical monocytes7,615 cells Tem/Temra cytotoxic T cells7,013 cells Tem/Trm cytotoxic T cells4,771 cells CD24 neutrophils2,333 cells Neutrophil-myeloid progenitor2,022 cells Early erythroid1,889 cells Tem/Effector helper T cells1,604 cells CRTAM+ gamma-delta T cells1,371 cells Memory B cells1,061 cells Non-classical monocytes961 cells Tcm/Naive cytotoxic T cells698 cells Regulatory T cells580 cells NAMPT neutrophils443 cells Megakaryocyte precursor397 cells Follicular helper T cells280 cells Age-associated B cells135 cells Megakaryocytes/platelets53 cells Small pre-B cells42 cells 10x 5' transcription profiling3,230 cells African American or Afro-Caribbean3,417 cells Hispanic or Latin American536 cells TSP14_BoneMarrow_NA_10X_1_12,593 cells TSP14_BoneMarrow_NA_10X_2_11,696 cells TSP2_BM_vertebralbody_10X_1_11,261 cells CZI-IA10471909+CZI-IA104720051,028 cells TSP11_BoneMarrow_NA_10X_1_1_LinEnriched987 cells CZI-IA10471910+CZI-IA10472006968 cells TSP14_BoneMarrow_NA_10X_2_1_5Prime793 cells TSP2_BM_vertebralbody_10X_2_1736 cells CZI-IA10244331+CZI-IA10244433703 cells TSP14_BoneMarrow_NA_10X_1_1_5Prime603 cells TSP11_BoneMarrow_NA_10X_1_1_LinDepleted469 cells CZI-IA10466284+CZI-IA10466380427 cells CZI-IA10466283+CZI-IA10466379403 cells CZI-IA10466281+CZI-IA10466377393 cells CZI-IA10466282+CZI-IA10466378367 cells TSP2_BM_vertebralbody_10X_2_2_5prime367 cells TSP11_BM_NA_SS2_B134009_B003956337 cells TSP2_BM_vertebralbody_10X_1_2_5prime334 cells TSP2_BM_vertebralbody_SS2_B113700_B133089301 cells TSP11_BM_NA_SS2_B133711_B003959263 cells TSP11_BM_NA_SS2_B133715_B003958241 cells TSP13_BM_vertebralbody_SS2_B133713_B134048_Endoth226 cells TSP11_BM_NA_SS2_B133717_B003957190 cells TSP2_BM_vertebralbody_SS2_B113709_B134460180 cells TSP13_BM_vertebralbody_SS2_B133720_B134046_HSC175 cells TSP2_BM_vertebralbody_SS2_B113698_B133341173 cells TSP14_BoneMarrow_NA_10X_4_1162 cells TSP13_BM_vertebralbody_SS2_B133712_B134041_Immune106 cells TSP2_BM_vertebralbody_SS2_B113699_B13309065 cells TSP13_BM_vertebralbody_SS2_B133719_B134047_Stromal28 cells TSP14_BoneMarrow_NA_10X_3_16 cells TSP13_BM_vertebralbody_SS2_B133719_B134047_Empty_1 cells 